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Allen and Greenough/ New Latin Grammar

Gender in the Fourth Declension

90. Most nouns of the Fourth Declension in -us are Masculine.

Exceptions: The following are Feminine: acus, anus, colus, domus, īdūs (plural), manus, nurus, porticus, quīnquātrūs (plural), socrus, tribus, with a few names of plants and trees. Also, rarely, penus, specus.

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